--- title: "Density Dependent IPMs" output: rmarkdown::html_vignette: toc: yes toc_depth: 3 vignette: > %\VignetteIndexEntry{Density Dependent IPMs} %\VignetteEngine{knitr::rmarkdown} %\VignetteEncoding{UTF-8} --- # Density dependent models Density dependent model classes are now implemented. This vignette will get more details shortly. For now, see the example below: ## Example of a simple, stochastic, kernel-resampled model with density dependence This example assumes that density dependence is modeled as a fixed effect in survival and recruit production models, and assumes there is no density dependence in growth or probability of reproducing models. The survival ($s(z, N)$/`s_yr`), growth ($G_{yr}(z',z)$/ `g_yr`), and number of recruit models ($r_{s,yr}(z, N)$/`r_s_yr`) have year-specific intercepts as well. The mathematical form for the IPM is below: 1. $n(z', t+1) = K_{yr}(z', z, N)n(z, t)dz$ 2. $N = \int_L^Un(z,t)dz$ 3. $K_{yr}(z', z, N) = P_{yr}(z', z, N) + F_{yr}(z', z, N)$ Here, $N$ represents the total population size. The kernel values fluctuate as a function of $N$ at each iteration of the model. The $P_{yr}(z', z, N)$ kernel is comprised of a density independent function for growth (Eq 6-7) and a density dependent function for survival (Eq 5). $f_G$ denotes a Gaussian probability density function: 4. $P(z', z, N) = s(z, N) * G(z', z)$ 5. $Logit(s(z, N)) = \alpha_s + \alpha_{s,yr} + \beta_s^z * z + \beta_s^{N} * N$ 6. $G(z', z, \theta) = f_G(z', \mu_{G,yr}(z), \sigma_G)$ 7. $\mu_{G,yr}(z) = \alpha_G + \alpha_{G,yr} + \beta_G^z * z$ The $F_{yr}(z',z, N)$ kernel is comprised of a density independent function for recruit size (Eq 10) and probability of reproducing (Eq 9), and a density dependent function for number of recruits produced by parents (Eq 11). $f_{r_d}$ denotes a Gaussian probability density function: 8. $F_{yr}(z', z, N) = r_r(z) * r_{s,yr}(z, N) + r_d(z')$ 9. $Logit(r_r(z)) = \alpha_{r_r} + \beta_{r_r}^z * z$ 10. $r_d(z') = f_{r_d}(z', \mu_{r_d}, \sigma_{r_d})$ 11. $Log(r_{s,yr}(z, N)) = \alpha_{r_s} + \alpha_{{r_s},yr} + \beta_{r_s}^z * z + \beta_{r_s}^N * N$ We'll simulate a 50 year time series using hypothetical parameter values. The fixed parameter values are created as with a density independent model. The difference is that we now have two more parameters: `s_dd`, and `r_s_dd`. These are the coefficients that correspond to $\beta_s^N$ and $\beta_{r_s}^N$, respectively. The chunk below initializes the data list object, which we name `params`. ```{r eval = FALSE} library(ipmr) data_list = list( s_int = 1.03, s_slope = 2.2, s_dd = -0.7, g_int = 8, g_slope = 0.92, sd_g = 0.9, r_r_int = 0.09, r_r_slope = 0.05, r_s_int = 0.1, r_s_slope = 0.005, r_s_dd = -0.03, mu_rd = 9, sd_rd = 2 ) # Now, simulate some random intercepts for growth, survival, and offspring production g_r_int <- rnorm(5, 0, 0.3) s_r_int <- rnorm(5, 0, 0.7) r_s_r_int <- rnorm(5, 0, 0.2) nms <- paste("r_", 1:5, sep = "") names(g_r_int) <- paste("g_", nms, sep = "") names(s_r_int) <- paste("s_", nms, sep = "") names(r_s_r_int) <- paste("r_s_", nms, sep = "") params <- c(data_list, g_r_int, s_r_int, r_s_r_int) ``` Next, we initialize the model using `init_ipm`. The difference is that the second argument is now changed to `"dd"` to denote that this is a density dependent model. ```{r eval = FALSE} dd_ipm <- init_ipm(sim_gen = "simple", di_dd = "dd", det_stoch = "stoch", kern_param = "kern") ``` Once we've done that, we're ready to begin specifying the kernel forms. One previously not mentioned aspect of `define_pop_state()` is that, in addition to defining initial conditions, 2 additional helper variables are generated: `n_stateVariable_t` and `n_stateVariable_t_1`. These can be used to reference the population states in vital rate and/or kernel expressions. These will look very similar to the ones we specified for density-independent models, except that we now include the term `s_dd * sum(n_size_t)` in the survival expression. `sum(n_size_t)` is the syntax `ipmr` uses to denote total population size. Further down, there is an example of how to use subsets of the trait distribution. ```{r eval = FALSE} dd_ipm <- define_kernel( proto_ipm = dd_ipm, name = "P_yr", formula = s_yr * g_yr, family = "CC", s_yr = plogis(s_int + s_r_yr + s_slope * size_1 + s_dd * sum(n_size_t)), g_yr = dnorm(size_2, g_mu_yr, sd_g), g_mu_yr = g_int + g_r_yr + g_slope * size_1, data_list = params, states = list(c("size")), uses_par_sets = TRUE, par_set_indices = list(yr = 1:5), evict_cor = TRUE, evict_fun = truncated_distributions("norm", "g_yr") ) ``` Other than the inclusion of the density dependent term in the survival expression, this should look quite similar to the density-independent kernel-resampled models from the Introduction vignette. We are now ready to continue defining the $F_{yr}(z',z,N)$ kernel. ```{r eval = FALSE} dd_ipm <- define_kernel( proto_ipm = dd_ipm, name = "F_yr", formula = r_r * r_s_yr * r_d, family = "CC", r_r = plogis(r_r_int + r_r_slope * size_1), r_s_yr = exp(r_s_int + r_s_r_yr + r_s_slope * size_1 + r_s_dd * sum(n_size_t)), r_d = dnorm(size_2, mu_rd, sd_rd), data_list = params, states = list(c("size")), uses_par_sets = TRUE, par_set_indices = list(yr = 1:5), evict_cor = TRUE, evict_fun = truncated_distributions("norm", "r_d") ) ``` Again, we've add the `f_s_dd * sum(n_size_t)` to the expression for `f_s_yr`, but otherwise, not much is different from how we've defined density independent models. The rest of the model definition process is unchanged. ```{r eval = FALSE} dd_ipm <- dd_ipm %>% define_impl( make_impl_args_list( kernel_names = c("P_yr", "F_yr"), int_rule = rep("midpoint", 2), state_start = rep("size", 2), state_end = rep("size", 2) ) ) %>% define_domains( size = c(0, 50, 200) ) %>% define_pop_state( n_size = runif(200) ) %>% make_ipm( iterate = TRUE, iterations = 50, kernel_seq = sample(1:5, 50, replace = TRUE) ) ``` `lambda` methods are defined for all density-dependent models as well. It is fairly straightforward to plot population sizes for these models by extracting the column sums of the arrays in `pop_state`. ```{r eval = FALSE} time_step_lams <- lambda(dd_ipm, type_lambda = "all") stoch_lam <- lambda(dd_ipm, type_lambda = "stochastic", burn_in = 0.15) pop_sizes <- colSums(dd_ipm$pop_state$n_size) plot(pop_sizes, type = "l") ```